Psammodes has been revised:
Female terminalia morphology and cladistic relations among Tok-Tok beetles (Tenebrionidae: Sepidiini)
Marcin J. Kamiński, Olivia M. Gearner, Marcin Raś, Elliot T. Hunsinger, Amelia L. Smith, et al. 2022
https://doi.org/10.1111/cla.12510 Cladistics 38: 623-648
Abstract:
Tok-tokkies are one of the most iconic lineages within Tenebrionidae. In addition to containing some of the largest darkling beetles, this tribe is recognized for its remarkable form of sexual communication known as substrate tapping. Nevertheless, the phylogenetic relationships within the group remain poorly understood. This study investigates the usefulness of female terminalia morphology for delimiting Sepidiini and reconstructing relationships among it.
Data on the structure of the ovipositors, genital tubes and spicula ventrali have been generated for >200 species representing 28 Pimeliinae tribes. This dataset was used in a comparative analysis at the subfamilial level, which resulted in recognition of several unique features of tok-tokkie terminalia.
Additionally, new features linking phenotypically challenging tribes also were recovered (Cryptochilini + Idisiini + Pimeliini).
Secondly, 23 characters linked to the structure of female terminalia were defined for tok-tok beetles. Cladistic analysis demonstrates the non-monophyletic nature of most of the recognized subtribes. The morphological dataset was analysed separately and in combination with available molecular data (CAD, Wg, cox1, cox2, 28S). All obtained topologies were largely congruent, supporting the following changes:
- Palpomodina Kamiński & Gearner subtr.n. is erected to accommodate the genera Namibomodes and Palpomodes;
- Argenticrinis and Bombocnodulus are transferred from Hypomelina to Molurina;
153 species and subspecies previously classified within Psammodes are distributed over three separate genera: Mariazofia, Piesomera & Psammodes.
Psammodes sklodowskae Kamiński & Gearner sp.n. is described. Preliminary investigation of the ovipositor of Mariazofia basuto (Koch) comb.n. was carried out with the application of microcomputed tomography, illuminating the muscular system as a reliable reference point for recognizing homologous elements in highly modified ovipositors.
-------------------------------------------------------------------------------------------
That has led to this proposed swap:
https://www.inaturalist.org/taxon_changes/113803
& this discussion:
https://www.inaturalist.org/flags/584700
So, in short, most of our IDs on iNat are out of date as Psammodes, but have not been fixed because doing so will change over 500 Mariazofia IDs (as intended by anyone using the local field guides) to Subtribe Molurina - which will require a monumental effort to get back to Mariazofia (basically hundreds of people will have to redo their IDs).
Please help us resolve this issue by tackling some of he problems below.
