following Morales-Jimenez, Alba Lucia, Liliana Cortés-Ortiz, and Anthony Di Fiore. "Phylogenetic relationships of Mesoamerican spider monkeys (Ateles geoffroyi): Molecular evidence suggests the need for a revised taxonomy." Molecular phylogenetics and evolution 82 (2015): 484-494.

sites.lsa.umich.edu/cortes-ortiz-lab/wp-content/uploads/sites/568/2019/06/Morales-Jimenez-et-al.-2015.pdf

mtDNA lumped yucatanensis into vellerosus