Tuolomne County is a medium-sized county that extends from the grasslands and chaparral at the eastern edge of the Central Valley, through the oak and Grey Pine woodlands of the Sierra foothills. From there, it extends into the coniferous forests and high alpine habitats. Much of the county is wilderness composed of the Stanislaus National Forest. It also holds the northern portion of Yosemite National Park (but none of the major tourist areas other than Tuolomne Meadow), the southern portion of Calaveras Big Trees State Park, and several large lakes and reservoirs on its western edge.

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