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I'm not sure of the exact paper - do you think POWO, BODATSA, and Solanaceae Source may have made a mistake?
Here is the original flag: https://www.inaturalist.org/flags/592817
I see. there is some info on it here: https://solanaceaesource.myspecies.info/content/solanum-humile
Do you know the paper where this was done?